In non-flowering seed plants, pollen germinates in the pollen chamber, located beneath the micropyle, underneath the integuments of the ovule.
2.
The pollen enters a pollen chamber close to the nucellus, and there it may wait for a year before it germinates and forms a pollen tube that grows through the wall of the megasporangium ( = nucellus ) where fertilisation takes place.
3.
These earliest ovules had the apical part of the nucellus exposed, from which there was a projection known as a lagenostome ( sometimes also called a salpinx ) that facilitated capture of the pollen and directed it down to the pollen chamber above the megagametophyte.
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