The roof of the mesopterygoid fossa, the opening behind the palate, is not perforated by sphenopalatine vacuities . " O . gorgasi " lacks an alisphenoid strut; in some other oryzomyines, this extension of the alisphenoid bone separates two openings in the skull, the masticatory buccinator foramen and the foramen ovale accessorium.
12.
In the first place, examination of embryos has revealed that the development of the braincase wall is essentially identical in therians and in'prototherians': the anterior lamina simply fuses with the alisphenoid in therians, and therefore the'prototherian'condition of the braincase wall is primitive for all mammals while the therian condition can be derived from it.
13.
Prototheria, on the other hand, was generally recognised as a subclass until quite recently, on the basis of an hypothesis which defined the group by two supposed synapomorphies : ( 1 ) formation of the side wall of the braincase from a bone called the anterior lamina, contrasting with the alisphenoid in therians; and ( 2 ) a linear alignment of molar cusps, contrasting with a triangular arrangement in therians.
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