It was discovered in 2011 by Julio C�sar Dalponte, and recognized as a new species in 2014 ( from the genus " Callicebus thomas " ).
12.
A pair of white-eared titis ( " P . donacophilus " ) entwining tails . Historically, titis were monogeneric and formed the genus " Callicebus"
13.
Together with Javier Garcia, he led an expedition in which they discovered and described a new species of titi monkey, the Caquet?titi ( " Callicebus caquetensis " ).
14.
Although the exact position of species within the " Callicebus " genus is debated, but they are treated as separate species in the latest edition of Mammal Species of the World.
15.
But whether the Lucifer titi, " Callicebus lucifer ", or the Colombian black-handed titi, " Callicebus medemi ", are good species from this complex is in doubt.
16.
But whether the Lucifer titi, " Callicebus lucifer ", or the Colombian black-handed titi, " Callicebus medemi ", are good species from this complex is in doubt.
17.
This extension to the ears is weak in " Callicebus torquatus torquatus ", a subspecies not confirmed for Colombia and different from the other subspecies which have white extending to the base of the ears.
18.
The "'collared titi "', " Callicebus torquatus ", is a species or a closely related complex of species of titi, a type of New World monkey, from South America.
19.
At the end of the 1980s the " Callicebus " genus was revised from the Hershkovitz concept of three species to thirteen neotropical species, with the collared titi, " Callicebus torquatus ", having four subspecies.
20.
At the end of the 1980s the " Callicebus " genus was revised from the Hershkovitz concept of three species to thirteen neotropical species, with the collared titi, " Callicebus torquatus ", having four subspecies.
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