The quadrate of " P . longirostris " is not very well preserved; it agees with " Odontopteryx " in a broad main shaft but is closer to " Osteodontornis " in the straight main shaft ridge and its upward-directed ventral articulation ridge's forward center.
12.
Its quadrate bone had a broad main shaft like in " Odontopteryx " which like in that genus bore a broad lateral ridge that was not, however, curved, but straight as in " Osteodontornis "; like in that genus, the forward center of the quadrate's ventral articulation ridge extended upwards.
13.
Its paroccipital process is not as markedly elongated back-and downwards as in the Ypresian ( Early Eocene ) " Dasornis " and " Odontopteryx " and seems to be in a more apomorphic condition, which would agree with a late Paleogene, possibly even ( like " Pelagornis " ) Neogene age for the holotype.
14.
The quadrate of the mysterious " Pseudodontornis longirostris " skull ( which some consider to belong in " Pelagornis " ) is not very well preserved; it agees with " Odontopteryx " in a broad main shaft and with " Osteodontornis " in the straight main shaft ridge and its upward-directed ventral articulation ridge's forward center.
15.
But the fossil femur measures only half as large as that of " Osteodontornis " ( or " P . longirostris ", for that matter ) . " " P . " stirtoni " was thus marginally larger than the Australian pelican ( " Pelecanus conspicillatus " ) of our time, or about the size of the small Paleogene pseudotooth bird " Odontopteryx toliapica ".
16.
"Odontopteryx " differed from " Pelagornis " ( a contemporary of " Osteodontornis " ) and agreed with " Dasornis " in having a deep and long handward-pointing pneumatic foramen in the fossa pneumotricipitalis of the humerus, a latissimus dorsi muscle attachment site on the humerus that consists of two distinct segments instead of a single long, and a large knob that extends along the ulna where the ligamentum collaterale ventrale attached.
17.
Further traits in which " Dasornis " agreed with " Odontopteryx " & ndash; and differed from " Pelagornis " ( a contemporary of " Osteodontornis " ) are a deep and long handward-pointing pneumatic foramen in the fossa pneumotricipitalis of the humerus, a latissimus dorsi muscle attachment site on the humerus that consists of two distinct segments instead of a single long, and a large knob that extends along the ulna where the ligamentum collaterale ventrale attached.
18.
But even though & ndash; due to the lack of better-preserved fossils & ndash; a close relationship between " Odontopteryx " and " Dasornis " cannot be excluded for sure either, it seems that the Neogene pseudotooth birds all derive from a large Paleogene form & ndash; such as " Dasornis " or ( if it is not actually identical with " Pelagornis " ) the mysterious " P . longirostris " & ndash; and that the smallish lineage became entirely extinct before the Neogene ( perhaps in the " Grande Coupure " ).
19.
One to five ( or perhaps more ) additional unnamed species are tentatively assigned to the present genus, mainly due to their size and / or forward-angled " teeth " : one smaller and one larger than " O . toliapica " and also from the Late Paleocene or Early Eocene of the Ouled Abdoun Basin in Morocco, one from the mid-Eocene of Uzbekistan, one from Middle Eocene strata of the Tepetate Formation from near El Cien ( Baja California Sur, Mexico ), and one from the Early Eocene of Virginia, USA . As regards the Moroccan fossils, however, the largest of the three " Odontopteryx "-like forms ( initially called " " Odontopteryx " n . sp . 2 " ) has provisionally been termed "'" Odontopteryx gigas " "'but may in fact be a " Dasornis ", while the smallest ( " " Odontopteryx " n . sp . 1 " ) has been considered a distinct genus ( as " Odontoptila inexpectata " ) but that name is both a " nomen nudum " and would in any case be a junior homonym of the geometer moth genus " Odontoptila " and thus unavailable for the bird.
20.
One to five ( or perhaps more ) additional unnamed species are tentatively assigned to the present genus, mainly due to their size and / or forward-angled " teeth " : one smaller and one larger than " O . toliapica " and also from the Late Paleocene or Early Eocene of the Ouled Abdoun Basin in Morocco, one from the mid-Eocene of Uzbekistan, one from Middle Eocene strata of the Tepetate Formation from near El Cien ( Baja California Sur, Mexico ), and one from the Early Eocene of Virginia, USA . As regards the Moroccan fossils, however, the largest of the three " Odontopteryx "-like forms ( initially called " " Odontopteryx " n . sp . 2 " ) has provisionally been termed "'" Odontopteryx gigas " "'but may in fact be a " Dasornis ", while the smallest ( " " Odontopteryx " n . sp . 1 " ) has been considered a distinct genus ( as " Odontoptila inexpectata " ) but that name is both a " nomen nudum " and would in any case be a junior homonym of the geometer moth genus " Odontoptila " and thus unavailable for the bird.
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