31. It produces separate sterile and fertile fronds, up to tall, and is a pioneer species on acidic screes. 32. It grows in sunny locations and a variety of soils, and functions as a pioneer species in ecological succession. 33. In a psammosere, the organisms closest to the sea will be pioneer species : environment, in particular the climate. 34. It is a pioneer species , colonising vacant land and forming mixed forests as other trees appear in its wake. 35. Unicellular floating algal plants such as diatoms are pioneer species of a bare water body, such as a pond. 36. "P . didactyla " is a common pioneer species on disturbed soils and nutrient poor grasslands in Western Europe. 37. This grass is found in many types of habitat, including disturbed areas where it acts as a pioneer species . 38. The new soil was re-colonized by pioneer species , which coexist alongside an established afforestation, forming a mosaic of habitats. 39. Pioneer species , which inhabit the plots of the forest destroyed by logging and fire, include sumac and black locusts.40. It is a pioneer species that thrives in poor soil, able to tolerate acidic and alkaline conditions and drought.