But by 1988 he had reduced Coptidoideae to a tribe within Isopyroideae, leaving five subfamilies, an arrangement he continued in his 1993 monograph, dividing the larger subfamilies into tribes, though by then " Paeonia " and " Glaucidium " were no longer considered to belong to Ranunculaceae . " Paeonia " was separated from Ranuculaceae and placed in its own family of Paeoniaceae ( order Saxifragales ) . other genera originally included in Ranunculaceae include " Circaeaster " which was placed in its own family Circaeasteraceae.
42.
But by 1988 he had reduced Coptidoideae to a tribe within Isopyroideae, leaving five subfamilies, an arrangement he continued in his 1993 monograph, dividing the larger subfamilies into tribes, though by then " Paeonia " and " Glaucidium " were no longer considered to belong to Ranunculaceae . " Paeonia " was separated from Ranuculaceae and placed in its own family of Paeoniaceae ( order Saxifragales ) . other genera originally included in Ranunculaceae include " Circaeaster " which was placed in its own family Circaeasteraceae.
43.
His work on the evolutionary biology of plants of interest to horticulture includes relationships within " Pelargonium " ( a collaboration with Mary Gibby of the Royal Botanic Garden Edinburgh ), evolutionary patterns within plant genera on islands, with emphasis on " Echium ", the phylogenetic history of " Drosera " and phylogenetic and taxonomic revision on " Actaea " and Ranunculaceae tribe Actaeae ( in collaboration with James Compton ), a phylogenetic evaluation of " Plectranthus " and " Solenostemon " ( coleus ) ( in collaboration with Alan Paton from the Royal Botanic Gardens, Kew ) and recently a review of " Cyclamen " phylogeny and classification ( in collaboration with James Compton and The Cyclamen Society ).
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