1. However, most modern groups of spiders have lost the cribellum . 2. The lack of a functional cribellum in araneoids is most likely synapomorphic. 3. They often have a calamistrum on metatarsus IV associated with a cribellum . 4. The cribellates retained the ancestral character, yet the cribellum was lost in the escribellates. 5. Placing Leptonetidae as a derived member of the Haplogynae would require independent evolution of the cribellum . 6. The ratio between calamistrum length and cribellum width varies greatly, however, even among related species. 7. They also have a calamistrum an apparatus of bristles used to comb the cribellate silk from the cribellum . 8. The length of a spider's calamistrum is always equal to or greater than the width of the cribellum . 9. Instead, to capture prey the spider uses its legs to comb webbing across its cribellum , a spiked plate near the spinnerets. 10. The Deinopoidea ( including the Uloboridae ), have a cribellum a flat, complex spinning plate from which the cribellate silk is released.