In the early 1920s, Howell isolated a water-soluble polysaccharide anticoagulant, which he also termed'heparin', although it was different from the previously discovered phosphatide preparations.
2.
McLean was a second-year medical student at Johns Hopkins University, and was working under the guidance of Howell investigating pro-coagulant preparations, when he isolated a fat-soluble phosphatide anticoagulant in canine liver tissue.
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