In different earthworm families, the typhlosole appears to have multiple origins.
2.
The mucous membranes of hagfishes have a primitive typhlosole in the form of permanent zigzag ridges.
3.
"Sparganophilus " species are characterised by a simple gut with no gizzard and no typhlosole.
4.
The lamprey intestine has a typhlosole that increases the inner surface like the spiral valve does in some jawed vertebrates.
5.
The spiral valve in the latter develops by twisting the whole gut, while the lamprey typhlosole is confined to the mucous membrane of the intestines.
6.
The Lumbricidae, for example, have a typhlosole which is an infolding of all layers of the intestine wall, whereas in some other families ( e . g.
7.
The Criodrilidae are characterised by holoic nephridia absent from anterior segments ( cf . " Pontodrilus " ), a simple gut with no gizzard and no typhlosole.
8.
"Criodrilidae " member species are characterised by holoic nephridia absent from anterior segments ( cf . " "'Pontodrilus " "'), a simple gut with no gizzard and no typhlosole.
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