The genera " Anancus ", " Morrillia ", " Paratetralophodon ", and " Tetralophodon ", but also Choerolophodontidae and Amebelodontidae were formerly classified as gomphotheres, but recent work recovers gomphotheres " sensu lato ", as paraphyletic, with tetralophodont gompotheres closely related to Elephantidae and amebelodonts and choerolophodonts more primitive than trilophodont gomphotheres.
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